The following are papers we have published related to our work on mountain lions in the Santa Cruz Mountains.  For a complete list of our lab’s publications, please visit



Smith, J.A., Thomas, A.C., Levi, T., Wang, Y. & Wilmers, C.C. 2018. Human activity reduces niche partitioning among three widespread mesocarnivores. Oikos 00:1-12. (PDF)

Tucker et al. 2018. Moving in the Anthropocene: Global reductions in terrestrial mammalian movements. Science 359: 466-469. (PDF)



Wang, Y., Smith, J.A. & Wilmers, C.C. 2017. Residential development alters behavior, movement, and energetics in an apex predator, the puma. Plos One 12:1-17. (PDF)

Bryce, C.M., Wilmers, C.C. & Williams, T.M. 2017. Energetics and evasion dynamics of large predators and prey: pumas vs. hounds. PeerJ:1-23. (PDF)

Smith, J. A., Suraci, J. P., Clinchy, M., Crawford, A., Roberts, D., Zanette, L. Y., & Wilmers, C. C. 2017. Fear of the human ‘super predator’ reduces feeding time in large carnivores. Proceedings of the Royal Society B 284:1-5. (PDF)

Allen, M. L., Wang, Y., & Wilmers, C. C. 2017. Exploring the adaptive significance of five types of puma (puma concolor) vocalizations. Canadian Field-Naturalist 130:289-294. (PDF)

Suraci, J. P., Clinchy, M., Mugerwa, B., Delsey, M., Macdonald, D. W., Smith, J. A., Wilmers, C. C., & Zanette, L. Y. 2017. A new automated behavioural response system to integrate playback experiments into camera trap studies. Methods in Ecology and Evolution 8:957-964. (PDF)



Allen, M.L., Gunther, M.S. & Wilmers, C.C. 2017. The scent of your enemy is my friend? The acquisition of large carnivore scent by a smaller carnivore. Journal of Ethology.(PDF)

Allen, M.L., Wilmers, C.C., Elbroch, L.M., Golla, J.M. & Wittmer, H.U. 2016. The importance of motivation, weapons and foul odors in driving encounter competition in carnivores. Ecology 97: 1905-1912. (PDF)

Smith, J.A., Wang, Y. & Wilmers, C.C. 2016. Spatial characteristics of residential development shift large carnivore prey habits. Journal of Wildlife Management.  (PDF)

Allen, M.L., Yovovich, V. & Wilmers, C.C. 2016. Evaluating the responses of a territorial solitary carnivore to potential mates and competitors. Scientific Reports 6:27257. (PDF)

Gray, M., Wilmers, C.C., Reed, S.E. & Merenlender, A.M. 2016. Landscape feature-based permeability models relate to puma occurrence. Landscape and Urban Planning 147: 50-58. (PDF)



Allen, M.L., Wittmer, H.U., Houghtaling, P., Smith, J., Elbroch, L.M. & Wilmers, C.C. 2015. The Role of Scent Marking in Mate Selection by Female Pumas (Puma concolor). Plos One: 1-12. (PDF)

Wilmers, C.C., Nickel, B., Bryce, C.M., Smith, J.A., Wheat, R.E. & Yovovich, V. 2015. The golden age of biologging:how animal-borne sensors are advancing the frontiers of ecology. Ecology 96: 1741-1753. (PDF)

Wang, Y., Allen, M.L. & Wilmers, C.C. 2015. Mesopredator spatial and temporal responses to large predators and human development in the Santa Cruz Mountains of California. Biological Conservation 190: 23-33. (PDF)

Allen, M.L., Elbroch, L.M., Wilmers, C.C. & Wittmer, H.U. 2015. The comparative effects of large carnivores on the acquisition of carrion by scavengers. American Naturalist 185: 822-833. (PDF)

Smith, J.A., Wang, Y. & Wilmers, C.C. 2015. Top carnivores increase their kill rates on prey as a response to human-induced fear Proceedings of the Royal Society B 282. (PDF)

Wang, Y., Nickel, B., Rutishauser, M., Bryce, C., Williams, T.M., Elkaim, G. & Wilmers, C.C. 2015. Movement, resting, and attack behaviors of wild pumas are revealed by tri-axial accelerometer measurements. Movement Ecology 3:2. (PDF)

Allen, M.L., Wallace, C.F. & Wilmers, C.C. 2015. Patterns in bobcat (Lynx rufus) scent marking and communication behaviors. Journal of Ethology 33: 9-14. (PDF)



Williams, T.M., Wolfe, L., Davis, T., Kendall, T., Richter, B., Wang, Y., Bryce, C., Elkaim, G.H. & Wilmers, C.C. 2014. Instantaneous energetics of puma kills reveal advantage of felid sneak attacks. Science 81: 81-85. (PDF)

Allen, M.L., Elbroch, M.L., Wilmers, C.C. & Wittmer, H.U. 2014. Trophic Facilitation or Limitation? Comparative Effects of Pumas and Black Bears on the Scavenger Community. Plos One 9. (PDF)

Allen, M.L., Wittmer, H.U. & Wilmers, C.C. 2014. Puma scrape and communication behaviors: understanding functional use and variation by sex and age. Behaviour 151: 819-840. (PDF)

Ripple, W.J., Estes, J.A., Beschta, R.L., Wilmers, C.C., Ritchie, E.G., Hebblewhite, M., Berger, J., Elmhagen, B., Letnic, M., Nelson, M.P., Schmitz, O.J., Smith, D.W., Wallach, A.D. & Wirsing, A.J. 2014. Status and ecological effects of the world’s largest carnivores. Science343:151. (PDF, Supplemental)



Wilmers, C.C., Wang, Y. , Nickel,  B., Houghtaling, P., Shakeri, Y., Allen, M. L., Kermish-Wells, J., Yovovich, V., & Williams, T. 2013. Scale dependent behavioral responses to human development by a large predator, the puma. Plos One 8:e60590. (PDF)